Mumble's son, Erik, is struggling to realize his talents in the Emperor Penguin world. Meanwhile, Mumble and his family and friends discover a new threat their home one that will take everyone working together to save them. Mumble the penguin has a problem: his son Erik, who is reluctant to dance, encounters The Mighty Sven, a penguin who can fly! Things get worse for Mumble when the world is shaken by powerful forces, causing him to brings together the penguin nations and their allies to set things right.

Into the world of the Emperor Penguins, who find their soul mates through song, a penguin is born who cannot sing. But he can tap dance something fierce! This is the story of a little penguin named Mumble who has a terrible singing voice and later discovers he has no Heartsong. However, Mumble has an astute talent for something that none of the penguins had ever seen before: tap dancing. Though Mumble's mom, Norma Jean, thinks this little habit is cute, his dad, Memphis, says it "just ain't penguin." Besides, they both know that, without a Heartsong, Mumble may never find true love. As fate would have it, his one friend, Gloria, happens to be the best singer around. Mumble and Gloria have a connection from the moment they hatch, but she struggles with his strange "hippity- hoppity" ways. Mumble is just too different--especially for Noah the Elder, the stern leader of Emperor Land, who ultimately casts him out of the community. Away from home for the first time, Mumble meets a posse of decidedly un-Emperor-like penguins--the Adelie Amigos. Led by Ramon, the Adelies instantly embrace Mumble's cool dance moves and invite him to party with...