Lady Tremaine gets her hands on the Fairy Godmother's wand, then turns back time to the day Cinderella tried on the glass slipper. She enlarges the slipper to fit one of the stepsisters, Anastasia, then erases Prince Charming's memories of meeting Cinderella. Now, Cinderella must foil Lady Tremaine's scheme and win back her prince.

Jacques and Gus tell the story of how Cinderella becomes a princess but loses touch with herself. The movie explores Cinderella's "happily ever after" life as a princess in three stories, with help from the Fairy Godmother. First, Cinderella's awkward first days at the palace, when she tried so hard to fit in that she forgot to be herself. Second, how Jaq felt so left out that he wished to be a human. Third, how Cinderella taught one of her nasty stepsisters how to smile which leads to her own true love.

When Cinderella's cruel stepmother prevents her from attending the Royal Ball, she gets some unexpected help from the lovable mice Gus and Jaq, and from her Fairy Godmother. In a far away, long ago kingdom, Cinderella is living happily with her mother and father until her mother dies. Cinderella's father remarries a cold, cruel woman who has two daughters, Drizella and Anastasia. When the father dies, Cinderella's wicked stepmother turns her into a virtual servant in her own house. Meanwhile, across town in the castle, the King determines that his son the Prince should find a suitable bride and provide him with a required number of grandchildren. So the King invites every eligible maiden in the kingdom to a fancy dress ball, where his son will be able to choose his bride. Cinderella has no suitable party dress for a ball, but her friends the mice, led by Jaques and Gus, and the birds lend a hand in making her one, a dress the evil stepsisters immediately tear apart on the evening of the ball. At this point, enter the Fairy Godmother, the pumpkin carriage, the royal ball, the stroke of midnight, the glass slipper, and the rest, as they say, is fairy tale ...

An alliance of evil-doers, led by Frieda, looks to take over Fairy Tale Land. But when Ella realizes her stepmother is out to ruin her storybook existence, she takes a dramatic turn and blossoms into the leader of the resistance effort. - As the story begins, an alliance of evil fairy tale-doers, led by Frieda, looks to take over Fairy Tale Land and take control of. But when Ella A.K.A. Cinderella realizes her own evil stepmother is out to ruin her storybook existence, she takes a dramatic turn and blossoms into the leader of the resistance effort.